At the end of the season, a fourth spore type, the teliospore, is formed.

The anthers, however, do not show outward signs of teliospore production, but are still effectively sterile because they tend to be small and immature.

Each teliospore undergoes karyogamy (fusion of nuclei) and meiosis to form four haploid spores called basidiospores.

Has a thick-walled resting spore (teliospore), known as the "brand" (burn) spore or chlamydospore.

The brothers were also able to make a connection between the red (urediniospore) and black (teliospore) spores as different stages within the same organism, but the rest of the stages remained unknown.

Pucciniomycetes develop no basidiocarp, karyogamy occurs in a thick-walled resting spore (teliospore), and meiosis occurs upon germination of teliospore.

It is a summer stage preceding the teleutospore, or winter stage.

Teliospore (sometimes called teleutospore) is the thick-walled resting spore of some fungi (rusts and smuts), from which the basidium arises.