In the anime, he serves as the primary villain.

They serve as primary villains of the first and second seasons.

He is the primary villain in the film.

The Ur-grue is revealed to be the primary villain of the story.

Melanie is known for being a primary villain, she has since become a responsible young woman.

The primary villains are mostly large fish or crustacean in form.

Queen Abront - the primary villain of the third series.

With the loss of two series regulars, producers had to look for a new primary villain for season nine.

He also appears as one of the primary villains from the sister show The Power Team.

She is known for being a primary villain.