Healthy, mature bulls in herds rarely fall victim to predators.
It was the smell of a mature bull, one ready to assume his place among the leaders of the herd.
Only mature bulls have large harems and breeding success peaks at about eight years of age.
The mature bulls kept a loose liaison with the breeding herd.
There were two mature bulls with the herd, but at the first threat they peeled away and disappeared into the forest.
She could only watch through two summers as the mature bulls passed her by.
This was the tenth day we had been hunting greater kudu and I had not seen a mature bull yet.
Fighting of mature bulls commences only after a special match, called "the Alternative".
All the elk were mature bulls, and the largest, with seven-point antlers, would have scored high in the record books.
A mature bull will produce 20 pounds or more of velvet annually.