The most dominant male will mate with females in the area.
He will "pretend" to be a female and take any opportunities presented to mate with willing females.
Males mate with several females, but have no part in raising the offspring.
Males may try to mate with females while calling or displaying.
Males choose to mate with females that are close to their own size.
During that time he may mate with other females, and add the eggs to his collection.
Males do not mate with breeding females that they are related to.
Therefore, sexual selection theory would suggest that a male should attempt to mate with several females.
Each male may meet and perhaps mate with several females in the course of moving through his range.
They had all formed a strong bond and are actively seeking to mate with females.