Several males may congregate at a lek site and these males are often closely related.
The males congregate regularly on damp ground.
Here, males and females congregate looking for mates.
The males also congregate in small groups but do not usually travel as far as the females.
In the early season, it is common that several males congregate around a newly emerged female.
During the mating season, males fight viciously, and may congregate around a single female.
Sub-adult males will congregate, often in association with an adult female.
Kin selection explains that related males congregate to form leks, as a way to attract females and increase inclusive fitness.
The mature males congregate at leks to breed, at which they display and are approached by females before mating with them.
During spawning, males congregate and create nests close together in colonies, and females visit to lay eggs.