"It was critical to get the landowners to agree to help us protect these young plants survive to produce seeds."
Most landowners along the Bull Run stream agreed to sell their rights for nominal fees.
The landowner agreed to further investigation, and a complete skull and tusk were found in 1974.
Douglas Mullen, the refuge's manager, said many landowners had agreed to sell and others were waiting to sell.
Greece's rich landowners, shipping magnates, doctors and dentists all agree to pay their taxes.
However, the landowner must agree to keep the infrastructure such as bridges and tunnels intact.
To mitigate the risk, the landowner agreed to inspect the tower every six months, and to take action upon discovery of further damage.
If the landowner agrees and the Federal Government picks up the costs, additional conservation measures can be imposed.
The landowner agreed as part of the settlement to address no further challenges to the project.
The landowner maintains actual ownership of the land, but agrees to certain development restrictions.