Algal blooms limit the sunlight available to bottom-dwelling organisms and cause wide swings in the amount of dissolved oxygen in the water.
He was the first to demonstrate that ceratioids were mesopelagic and bathypelagic lifeforms rather than bottom-dwelling organisms.
Juvenile halibut feed on small crustaceans and other bottom-dwelling organisms.
Federal scientists defend the tests, saying that samples that fail are clearly toxic to bottom-dwelling organisms.
Any such flows must have enveloped free-swimming as well as bottom-dwelling organisms.
Benthic refers to "bottom-dwelling" organisms that are known to exhibit unusual characteristics not normally seen in shallow-dwelling creatures.
But because many bottom-dwelling organisms - starfish and lobsters, for example - cannot move quickly, they may die in the oxygen-depleted lower layers.
They also include some bottom-dwelling (benthic) organisms in their diets.
No catastrophe was seen in surface ecosystems, yet bottom-dwelling organisms in the deep ocean experienced a major extinction.
Its diet includes benthic (bottom-dwelling) organisms like mollusks and worms.