Besides humans, no other creature penetrates the Northwest so completely.

It is feasible because there are no animal reservoirs besides humans and pigs.

"You mean they've found intelligent beings out there besides humans?"

I forgot that other things besides humans might fall into the pits.

Besides humans and dogs, the disease can be contracted by other animals, including horses.

Besides humans, they are the only free-living primates in Europe.

Most of them hadn't encountered many aliens before, at least none besides humans and Vulcans, who looked more or less like themselves.

There is another suitable sentient being besides humans in the colony.

"What creature besides humans would haunt this empty land, without a plant or animal to eat?"

Besides humans, predators that might thin the herds are few in this area.