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Like all cephalopods, the blood of the nautilus contains hemocyanin, which is blue in its oxygenated state.
Hemerythrin and myohemerythrin are essentially colorless when deoxygenated, but turn a violet-pink in the oxygenated state.
When under a low oxygenated state (hypoxia) the respiratory network responds by transitioning into an augmentation followed by a depression phase, controlled in the pre-BötC.
This has destabilized the T-state (the deoxygenated state of hemoglobin), which has led to a higher affinity for being in the R-state (oxygenated state of hemoglobin).
In his later experimental work Lemberg intensively studied, with Professor Ron Williams of Canada and Dr Marion Gilmour of the USA, the ferric state and the oxygenated state (discovered by Okunuki) of solubilized cytochrome oxidase.