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"And that is not to be confused with the quite different word, nucular, meaning 'of or relating to a nucule.' "
The fructifications for sexual reproduction are globule or antheridium (male) and nucule or archegonium (female).
Arguing by analogy, Elster suggests that "Molecular comes from molecule, and particular comes from particle, but there is no nucule to support nucular."
The references consulted do not agree as to whether the fruit is a nucule or achene, the difference between the two ultimately depends on the size of the pericarp and the extent of its lignification.
A large number of fossils of this family have been recorded from the Lower Cretaceous (98-113 million years ago, Platanocarpus), the examples from that time had very small pollen (8-10 μm) and a developed perianth and they lacked hairs at the base of the nucule.