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However, this is not thought to be for breeding, as the case in anadromous and catadromous species.
A catadromous species, it is tolerant of high and low salinities.
Finally, dams fragment river systems, isolating previously continuous populations, and preventing the migrations of anadromous and catadromous species.
Notwithstanding the geographical area of application of the agreement, Article 13 provides that the Parties may agree to extend cooperation on the management of anadromous and catadromous species, but excludes species which live their entire life cycle in internal waters.