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The bass's C pedal, becomes the tonic, when the key changes to either the aeolian or dorian modes.
This gives a dorian scale between 4 draw and 8 draw, though once again bends and overblows give players a variety of options.
Using this rhythmic phrasing, the dorian line played on the violins and the aeolian woodwind line are differentiated rhythmically, as well as tonally.
The horn call motif fluctuates from major to minor, outlining the tonal ambiguity, moving between the mixolydian and dorian modes, which becomes a characteristic of the movement.
A more advanced harmonic mixing theory has been proposed which accounts for the various modes as well (aeolian, ionian, lydian, mixolydian, dorian and phrygian).
The chord on the fifth scale degree may be major, V7, or minor, v7, in which case it fits a dorian scale along with the minor i7 and iv7 chords, creating a modal feeling.
"Cluck Old Hen" is a popular Appalachian fiddle and banjo tune in the mixolydian or dorian mode (as in the score below which is in A dorian).